:root{--bg:#f8f7f4;--surface:#ffffff;--nav-bg:#ffffff;--nav-text:#1c1c1c;--text-primary:#1c1c1c;--text-body:#3a3a3a;--text-muted:#7a7870;--accent-warm:#8b4513;--rule:#d8d5cf;--max-w:920px;--nav-h:62px;--font-serif:'Manrope', system-ui, sans-serif;--font-sans:'Poppins', system-ui, sans-serif;--font-mono:'IBM Plex Mono', 'Courier New', monospace}*,*::before,*::after{box-sizing:border-box;margin:0;padding:0}html{font-size:16px;scroll-behavior:smooth}body{font-family:var(--font-sans);font-weight:400;background:var(--bg);color:var(--text-body);line-height:1.75}img{display:block;max-width:100%;height:auto}a{color:inherit;text-decoration:none}.project-body a,.contact-body a,.page-intro a{text-decoration:underline;text-decoration-color:var(--text-muted);text-underline-offset:3px;text-decoration-thickness:1px;transition:text-decoration-color .2s}.project-body a:hover,.contact-body a:hover,.page-intro a:hover{text-decoration-color:var(--text-primary)}.site-nav{position:fixed;top:0;left:0;right:0;height:var(--nav-h);background:var(--nav-bg);display:flex;align-items:center;padding:0 2.5rem;z-index:100;box-shadow:0 1px 4px rgba(0,0,0,8%)}.nav-logo img{height:42px}.nav-logo-text{font-family:var(--font-serif);color:var(--nav-text);font-size:1.1rem}.nav-logo,.nav-logo-text{margin-right:auto}.nav-links{display:flex;gap:2rem;list-style:none}.nav-links a{font-size:.78rem;letter-spacing:.12em;text-transform:uppercase;color:rgba(28,28,28,.82);padding-bottom:2px;border-bottom:1px solid transparent;transition:color .2s,border-color .2s}.nav-links a:hover,.nav-links a.active{color:var(--nav-text);border-bottom-color:rgba(28,28,28,.35)}.nav-toggle{display:none;flex-direction:column;gap:5px;background:0 0;border:none;cursor:pointer;padding:4px}.nav-toggle span{display:block;width:22px;height:1.5px;background:rgba(28,28,28,.65);transition:transform .2s}main{padding-top:var(--nav-h)}.section-rule{border:none;border-top:1px solid var(--rule);max-width:var(--max-w);margin:0 auto}.section-label{font-size:.72rem;letter-spacing:.18em;text-transform:uppercase;color:var(--text-muted);margin-bottom:2.5rem;display:flex;align-items:center;gap:.8rem}.section-label::after{content:'';flex:1;height:1px;background:var(--rule)}.hero{max-width:var(--max-w);margin:0 auto;padding:6rem 2rem 3.5rem}.hero-intro{display:flex;align-items:flex-start;gap:3.5rem;margin-bottom:2rem}.hero h1{font-family:var(--font-serif);font-size:clamp(3.4rem,7vw,5.8rem);font-weight:800;color:var(--text-primary);line-height:1;letter-spacing:-.03em;margin:0;flex-shrink:0}.hero-sub{font-size:1.08rem;line-height:1.85;color:var(--text-body);padding-top:.6rem}.hero-sub p+p{margin-top:.7em}.hero-sub a{border-bottom:1px solid var(--rule);transition:border-color .2s}.hero-sub a:hover{border-color:var(--text-primary)}.listing-gallery{display:grid;grid-template-columns:repeat(4,1fr);gap:3px;margin-top:1.5rem}.listing-gallery figure{height:210px;overflow:hidden;background:var(--bg);margin:0}.listing-gallery figure img{width:100%;height:100%;object-fit:contain;display:block}.image-strip{display:grid;grid-template-columns:repeat(3,1fr);gap:3px;margin-top:3rem}.image-strip figure{aspect-ratio:4/3;overflow:hidden;background:#111}.image-strip figure img{width:100%;height:100%;object-fit:cover;opacity:.9;transition:transform .5s ease,opacity .3s}.image-strip figure:hover img{transform:scale(1.03);opacity:1}.section{max-width:var(--max-w);margin:0 auto;padding:5rem 2rem}.interest-block+.interest-block{margin-top:4rem}.interest-title{font-family:var(--font-serif);font-size:1.75rem;font-weight:700;color:var(--text-primary);margin-bottom:1.2rem;letter-spacing:-.02em}.interest-body,.section p{font-size:1rem;line-height:1.85;max-width:720px}.interest-body+.interest-body{margin-top:1em}.interest-block p{font-size:1rem;line-height:1.85;max-width:720px}.interest-block p+p{margin-top:1em}.interest-block em{font-style:italic}blockquote{border-left:2px solid var(--accent-warm);padding-left:1.4rem;margin:1.8rem 0;color:var(--text-muted);font-style:italic;font-size:.93rem;line-height:1.8;max-width:640px}.page-header{max-width:var(--max-w);margin:0 auto;padding:5rem 2rem 2.5rem;border-bottom:1px solid var(--rule)}.page-header h1{font-family:var(--font-serif);font-size:2.8rem;font-weight:700;color:var(--text-primary);letter-spacing:-.025em;margin-bottom:.9rem}.page-intro{font-size:1rem;max-width:620px;line-height:1.8}.page-intro h3{font-family:var(--font-serif);font-size:1.1rem;font-weight:600;color:var(--text-primary);margin-top:2.2em;margin-bottom:.4em}.page-intro p:last-child{margin-top:2em}.project-grid{max-width:var(--max-w);margin:0 auto;padding:3rem 2rem 5rem;display:grid;grid-template-columns:repeat(3,1fr);gap:1.5rem}.project-card{background:var(--surface);border:1px solid var(--rule);overflow:hidden;transition:border-color .2s,box-shadow .2s}.project-card:hover{border-color:#aaa9a4;box-shadow:0 4px 20px rgba(0,0,0,7%)}.project-card-link{display:block}.project-card-img{aspect-ratio:3/4;overflow:hidden;background:#0f0f0f}.project-card-img img{width:100%;height:100%;object-fit:cover;opacity:.88;transition:transform .45s ease,opacity .3s}.project-card:hover .project-card-img img{transform:scale(1.04);opacity:1}.project-card-body{padding:1.1rem 1.2rem 1.3rem}.project-card-tag{font-size:.67rem;letter-spacing:.14em;text-transform:uppercase;color:var(--text-muted);margin-bottom:.4rem}.project-card-title{font-family:var(--font-serif);font-size:1.05rem;font-weight:600;color:var(--text-primary);line-height:1.35}.project-hero{width:100%;max-height:480px;overflow:hidden;background:#0a0a0a}.project-hero img{width:100%;height:480px;object-fit:cover;opacity:.88}.project-content{max-width:var(--max-w);margin:0 auto;padding:3.5rem 2rem 5rem}.project-content h1{font-family:var(--font-serif);font-size:2.3rem;font-weight:700;color:var(--text-primary);letter-spacing:-.025em;margin-bottom:.5rem;max-width:700px;line-height:1.2}.project-content-tag{font-size:.7rem;letter-spacing:.14em;text-transform:uppercase;color:var(--text-muted);margin-bottom:2rem}.project-body{max-width:680px}.project-body p{font-size:1rem;line-height:1.85}.project-body p+p{margin-top:1.1em}.project-body h2{font-family:var(--font-serif);font-size:1.4rem;font-weight:600;color:var(--text-primary);margin:2.8rem 0 1rem;letter-spacing:-.01em}.project-body h2:first-child{margin-top:0}.project-body h3{font-family:var(--font-serif);font-size:1.15rem;font-weight:600;color:var(--text-primary);margin:2.2rem 0 .6rem;letter-spacing:-.005em}.project-body hr{border:none;border-top:1px solid var(--rule);margin:2.5rem 0}.project-body ul,.project-body ol{margin:1em 0 1em 1.5em;font-size:.95rem}.project-body li+li{margin-top:.3em}.image-gallery{display:grid;grid-template-columns:repeat(2,1fr);gap:3px;margin:3rem 0}.image-gallery figure{aspect-ratio:16/10;overflow:hidden;background:#0f0f0f}.image-gallery figure:first-child{grid-column:span 2;aspect-ratio:21/9}.image-gallery figure img{width:100%;height:100%;object-fit:cover;opacity:.9;transition:opacity .3s}.image-gallery figure:hover img{opacity:1}.project-nav{display:flex;justify-content:space-between;border-top:1px solid var(--rule);padding-top:2rem;margin-top:3rem}.project-nav-item{display:flex;flex-direction:column;gap:.3rem;max-width:45%}.project-nav-item span{font-size:.7rem;letter-spacing:.12em;text-transform:uppercase;color:var(--text-muted)}.project-nav-item a{font-family:var(--font-serif);font-size:1rem;color:var(--text-primary);border-bottom:1px solid transparent;transition:border-color .2s;line-height:1.3}.project-nav-item a:hover{border-color:var(--rule)}.project-nav-item.next{text-align:right}.callout{border-left:3px solid var(--accent-warm);background:#f3ede6;padding:1.4rem 1.6rem;margin:2.5rem 0;max-width:720px;border-radius:0 2px 2px 0}.callout p{font-size:.93rem;line-height:1.8;color:var(--text-body);margin:0}.callout p+p{margin-top:.8em}.callout strong{font-weight:600;color:var(--text-primary)}.project-figure{margin:2.5rem 0;max-width:720px}.project-figure img{width:100%;border:1px solid var(--rule);display:block}.project-figure figcaption{font-size:.82rem;line-height:1.65;color:var(--text-muted);margin-top:.6rem;padding-top:.5rem;border-top:1px solid var(--rule)}.project-figure figcaption strong{color:var(--text-body)}.project-figure figcaption sub,.project-figure figcaption sup{font-size:.75em}.project-body hr~p,.project-body p:has(>a[href*=biorxiv]){font-size:.88rem;color:var(--text-muted);line-height:1.7}.contact-body p{font-size:1rem;line-height:1.85;color:var(--text-body)}.contact-body p+p{margin-top:.9em}.contact-body h2{font-family:var(--font-serif);font-size:1.4rem;font-weight:600;color:var(--text-primary);letter-spacing:-.01em;margin:3rem 0 1rem}.contact-gap{clear:both;height:2.2rem}.contact-profile-img{float:left;width:155px;margin:.25rem 2.5rem 1rem 0;border-radius:4px;display:block}.contact-body::after{content:'';display:table;clear:both}@media(max-width:768px){.contact-profile-img{float:none;width:120px;margin:0 0 1.5rem}}.contact-page{max-width:var(--max-w);margin:0 auto;padding:6rem 2rem 5rem}.contact-grid{display:grid;grid-template-columns:repeat(2,1fr);gap:3rem 4rem;max-width:680px;margin-top:3rem}.contact-item-label{font-size:.7rem;letter-spacing:.16em;text-transform:uppercase;color:var(--text-muted);margin-bottom:.6rem}.contact-item a{font-size:.93rem;border-bottom:1px solid var(--rule);padding-bottom:1px;transition:border-color .2s}.contact-item a:hover{border-color:var(--text-primary)}.contact-item address{font-style:normal;font-size:.9rem;line-height:1.7;color:var(--text-body)}.contact-orcid{font-family:var(--font-mono);font-size:.82rem;color:var(--text-muted)}.contact-profiles{display:flex;flex-direction:column;gap:.5rem}.contact-profiles a{font-size:.9rem;display:flex;align-items:center;gap:.5rem;width:fit-content;border-bottom:1px solid var(--rule);padding-bottom:1px;transition:border-color .2s}.contact-profiles a:hover{border-color:var(--text-primary)}.site-footer{background:var(--nav-bg);color:rgba(28,28,28,.45);border-top:1px solid var(--rule);padding:2.2rem 2.5rem;display:flex;align-items:center;justify-content:space-between;font-size:.78rem;letter-spacing:.04em}.footer-name{display:block;font-family:var(--font-serif);font-size:.9rem;color:rgba(28,28,28,.85);margin-bottom:.25rem}.footer-affil{color:rgba(28,28,28,.6)}.footer-social{display:flex;gap:1.2rem}.footer-social a{color:rgba(28,28,28,.5);font-size:1rem;transition:color .2s}.footer-social a:hover{color:rgba(28,28,28,.85)}@media(max-width:768px){.site-nav{padding:0 1.5rem}.nav-links{display:none;position:absolute;top:var(--nav-h);left:0;right:0;background:var(--nav-bg);flex-direction:column;align-items:center;padding:2rem 1.5rem 2.5rem;gap:0;border-top:1px solid var(--rule);box-shadow:0 4px 8px rgba(0,0,0,6%)}.nav-links.open{display:flex}.nav-links li{width:100%;text-align:center}.nav-links li a{display:block;font-size:1rem;letter-spacing:.1em;padding:1rem 0;border-bottom:1px solid var(--rule)}.nav-links li:first-child a{border-top:1px solid var(--rule)}.nav-toggle{display:flex}.hero{padding:4rem 1.5rem 2.5rem}.hero-intro{flex-direction:column;gap:1.2rem}.hero-sub{padding-top:0}.listing-gallery{grid-template-columns:repeat(2,1fr)}.image-strip{grid-template-columns:1fr}.project-grid{grid-template-columns:1fr;gap:1rem;padding:2rem 1.5rem}.image-gallery{grid-template-columns:1fr}.image-gallery figure:first-child{grid-column:span 1;aspect-ratio:4/3}.section,.page-header,.project-content,.contact-page{padding-left:1.5rem;padding-right:1.5rem}.contact-grid{grid-template-columns:1fr;gap:2rem}.site-footer{flex-direction:column;gap:1.2rem;text-align:center}}@media(max-width:540px){.project-grid{grid-template-columns:1fr}}